ZOO keepers are mourning after one of the country's oldest African penguins died at the age of 37.
Pat, who lived at Living Coasts near Torquay and was thought to be the second-oldest penguin in Europe, was put down after suffering advanced arthritis.
Staff said she was put down because "her quality of life had declined".
A spokesman said: "Pat had a terrific innings thanks to good husbandry, good vet care, good food and freedom from predators. But her quality of life had declined because of arthritis and the only thing to do was to put her to sleep."